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Pyramidal cell types and 5-HT 2A receptors are essential for psilocybin’s lasting drug action

Ling-Xiao Shao, Clara Liao, Pasha A Davoudian, Neil K Savalia, Quan Jiang, Cassandra Wojtasiewicz, Diran Tan, Jack D Nothnagel, Rong-Jian Liu, Samuel C Woodburn, Olesia M Bilash, Hail Kim, Alicia Che, Alex C Kwan

bioRxiv (Cold Spring Harbor Laboratory) November 3, 2024 preprint DOI: 10.1101/2024.11.02.621692 via OpenAlex

Summary

AI-generated from the abstract

A single dose of psilocybin increased the density of dendritic spines in both subcortical-projecting pyramidal tract (PT) and intratelencephalic (IT) cell types in the mouse medial frontal cortex. Silencing PT neurons eliminated psilocybin's ability to ameliorate stress-related phenotypes, whereas silencing IT neurons had no detectable effect. In PT neurons only, psilocybin boosted synaptic calcium transients and elevated firing rates acutely after administration. Targeted knockout of 5-HT2A receptors abolished psilocybin's effects on stress-related behavior and structural plasticity. These results identify a pyramidal cell type and the 5-HT2A receptor in the medial frontal cortex as essential for psilocybin's long-term drug action.

Study at a glance

Characteristics In vivo optical imaging, chemogenetic perturbation, and cell type-specific electrophysiology
Population Mice
Intervention Psilocybin
Dose a single dose
Topics Depression Psilocybin Serotonin
Keywords Neural networks Neurobiology Mood disorders Psychiatric treatment Dendritic spines
Key finding Silencing PT neurons, but not IT neurons, eliminates psilocybin's ability to ameliorate stress-related phenotypes, and targeted knockout of 5-HT2A receptors abolishes psilocybin's effects on stress-related behavior and structural plasticity.

Abstract

Abstract Psilocybin is a serotonergic psychedelic with therapeutic potential for treating mental illnesses 1–4 . At the cellular level, psychedelics induce structural neural plasticity 5,6 , exemplified by the drug-evoked growth and remodeling of dendritic spines in cortical pyramidal cells 7–9 . A key question is how these cellular modifications map onto cell type-specific circuits to produce psychedelics’ behavioral actions 10 . Here, we use in vivo optical imaging, chemogenetic perturbation, and cell type-specific electrophysiology to investigate the impact of psilocybin on the two main types of pyramidal cells in the mouse medial frontal cortex. We find that a single dose of psilocybin increased the density of dendritic spines in both the subcortical-projecting, pyramidal tract (PT) and intratelencephalic (IT) cell types. Behaviorally, silencing the PT neurons eliminates psilocybin’s ability to ameliorate stress-related phenotypes, whereas silencing IT neurons has no detectable effect. In PT neurons only, psilocybin boosts synaptic calcium transients and elevates firing rates acutely after administration. Targeted knockout of 5-HT 2A receptors abolishes psilocybin’s effects on stress-related behavior and structural plasticity. Collectively these results identify a pyramidal cell type and the 5-HT 2A receptor in the medial frontal cortex as playing essential roles for psilocybin’s long-term drug action.

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